Current and past versions of FTOL available for download

About the data

Tree format

FTOL is available in several flavors:

All trees are rooted on algae (Zygnema) and include outgroup taxa, but Zygnema has been pruned from the dated trees. Node labels are bootstrap support values (%) measured with ultra-fast bootstrap (Hoang et al. 2018).

All trees are in Newick format.

Other data files

The tree files are provided as part of a data archive (zip or tar.gz format).

Other files in the archive include DNA sequence alignments, GenBank accession numbers, and fossil calibration points.

All files are described in the README.txt file contained within the archive.

ftolr R pacakge

For reading the data directly into R with more options (rooting, inclusion of outgroup, selection of only part of the alignment, etc.), use the ftolr R package.

Current version: v1.1.0

Built with data from GenBank release 249 (cutoff date 2022-04-15).

Past versions


Cutoff date 2021-12-31. Does not include algae and was rooted on bryophytes.


Hosted on FigShare and lacks some of the tree formats above due to changes in methodology.

Hoang, Diep Thi, Olga Chernomor, Arndt von Haeseler, Bui Quang Minh, and Le Sy Vinh. 2018. “UFBoot2: Improving the Ultrafast Bootstrap Approximation.” Molecular Biology and Evolution 35 (2): 518–22.